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<div>This variety have been recorded only in the department of Valle del Cauca, in humid forests, between 1900 and 2000 m. </div><div> Additional specimens examined </div><div><b>26.<i> Begonia solaniflora</i> </b>Jara. 2017. Phytotaxa 321(2): 209. Colombia: Santander, Suaita, corregimiento San José de Suaita, entre Marbella y La Meseta. 6.254°N, 73.4231°W, 1640 m, A. Jara & J. Luna 2568, 17 Dic 2013 (Holotype: ANDES!; Isotype: COL!).</div><div></div><div>Plant perennial, initially erect and becoming prostrate with age, up to 70 cm high. Internodes 6.7–31.3 (65.4) mm long, up to 2.5 mm across, reddish with some parts green, glabrous, nodes thickened. Stipules early caducous, 9.7−10.6 × 2.3−3.8 mm, lanceolate, glabrous, light green to hyaline, membranous, weakly keeled, glabrous, margin entire, apex acute to obtuse and aristate, ariste ca. 0.7 mm. Petioles 3.3–10.2 × 0.8–1.1 mm; narrowly canaliculate, with some bristles near the junction with the blade. Leaves straight, 4.1−8.5 × 1.4−2.5 mm, elliptic to ovate, slightly asymmetric, firmly membranous; base acute; apex acute; margin serrate with a hair between each pair of teeth; venation pinnate, with 3–6 major lateral veins on each side; tertiary veins generally terminating at the margin. Inflorescence axillary, unisexual. Staminate inflorescence: reduced monochasium; peduncle 6.1−9.2 mm long, glabrous; bracteoles 5–7, the most external up to 8.1 × 5.1 mm, the others centripetal contorts gradually smaller, hyaline to greenish, from broadly ovate the outer ones to elliptic the inner, margin entire, apex obtuse, reticulations evident. Staminate flowers: 5−7, pedicels 6.4−6.8 mm long; tepals 4, glabrous, margin entire, apex obtuse, outer 2 ovate, whitish, with some reddish veins parallels, 6.4–8.1 × 3.0-3.2, inner 2, elliptic, whitish, 6.5 × 1.3 mm; stamens 5–8, disposed upon an axis of ca. 0.9 mm, anthers 2-locular, opening internally, compressed, oblong-lanceolate, ca. 3.3 × 0.4 mm, thecae in contact among them by the inner side forming a staminal spindle-shape structure, connective ca. 0.7 mm, yellow. Pistillate inflorescence: flowers solitary, peduncle 5.1–8.4, glabrous; bracts 2, broadly ovate, 6.5–7.2 × 4.5–4.9 mm, hyaline, margin entire, apex apiculate. Pistillate flowers: sessile, tepals 5, margin entire; outer 2 tepals ovate, 9.3–14.4 × 1.9–3.0 mm, whitish, with reddish variegation, apex rounded to truncate; inner 3 tepals oblong-lanceolate, 10.3 × 0.9 mm, whitish, apex obtuse: ovary trigonous, 2.9 × 6.2 mm (horns excluded), glabrous; placentation simple, ovuliferous on both sides; styles 3, simple or once divided close to the base, 7.5–8.6 mm long, yellow; stigmatic tissue covering ca. 2.5 mm, all around the style tip. Fruit obconic, 9.1–9.5 × 20.8–24.4 mm (horns included), glabrous, horns flattened, apex acute, column absent.</div><div></div><h3 data-label="764335" class="ltx_title_subsubsection">Habitat and distribution</h3><div>Moist tropical montane forest at 1460–1800 m, growing near waterfalls and streams in semi-open or shaded places. Southern region of the department of Santander (Colombia) between the Chicamocha and Suarez Rivers.</div><div></div><h3 data-label="388837" class="ltx_title_subsubsection"> Additional specimens examined </h3><div>COLOMBIA: Santander, Suaita, corregimiento San José de Suaita, entre Marbella y La Meseta, 6°09’24’’N, 73°24’51’’W, ca. 1700 m. 17 Dic 2013, <i>A. Jara & J. Luna 2563</i> (ANDES), 2564 (ANDES), 2565 (ANDES), 2567</div><div> (ANDES); 2568(ANDES, HUA); 2569 (ANDES); 2570 (ANDES, COL, HUA); vereda Campo Anibal, 6°0.8’57.2’’–6°0.9’8.0’’N–73°24’54.5’’–73°24’33.3’’W, 1740–1810 m, 26 Mar 2004, Betancur et al. 10693(COL); Oiba, Cascadas de Chaguatá, 6°15’0’’N, 73°17’28’’W, 1460 m, 18 Dic 2013, Jara & Luna 2578 (ANDES, COL); 2579 (ANDES);2580 (UIS); 2581 (COL); 2582 (UIS); 2583 (COL); 2584 (UIS); 2585 (ANDES); 2586 (CUVC); 2587 (JAUM); 2588 (UDBC); 2589 (ANDES).</div><div></div><div><b>27. B<i>egonia suaviola </i></b>Jara (XX, 2016). Type:—COLOMBIA. Valle del Cauca: municipio de El Cairo, vereda El Brillante, reserva natural Cerro El Inglés, 2217–2328 m, 27 September 2013, <i>Jara, A. & Avila, F. 2447</i> (holotype ANDES!, isotypes CUVC!, HUA!, E! (to be distributed)). </div><div> </div><div>Erect herb, 20–40 cm tall. Stem 1.4–2.5 mm in diameter, surface villous with hairs ca. 1.3 mm long. Stipules persistent, oblong to ovate, asymmetrical, 4.6–13.0 × 2.0–4.6 mm, with some hairs, coriaceous, light green to whitish, margin entire, apex acute, with a terminal seta of ca. 1.0 mm long. Petiole 1.1–9.1 mm long, densely villous. Leaf lamina straight, oblong to elliptic, asymmetrical, 3.4–8.7 × 0.9–3.1 cm (2.4–4.1 length/width ratio), margin serrulate, base slightly oblique, apex long acuminate, strigose above, villous below, veins pinnate, with 6–10 pairs of veins. Inflorescences: monochasial cymes with very short internodes, bisexual or consisting only of staminate flowers, protrandrous; peduncle 15.5–38.4 mm long. Bracts caducous, 3.8–11.1 × 0.8–5.3 mm, elliptic to lanceolate; apex acute; margin entire. Secondary bracts similar to bracts. Staminate flowers: pedicels 4.0–10.4 mm long, villous; perianth campanulate with two perianth series, outer tepals 2 fused to ¼ their total length, obovate, 7.0–13.6 × 4.2–10.1 mm, emarginate to the apex, with a few villi at the base, orange-red, inner tepals 2, free, flavelate, apex rounded to emarginate, slightly longer than the stamens, ca. 3.1 × 3.8 mm, pale red, glabrous; stamens with short filaments ca. 0.3 mm long, glabrous, without torous, anthers oblong, ca. 2.2 × 0.7 mm, with connectives not pronounced. Pistillate flowers: pedicels 0.6– 3.4(7) mm long; bracteoles wanting; perianth campanulate with two series, outer tepals 2–3 fused ca. ¾ of their length, obovate, 15.5–19.5 × 11–14.4 mm, glabrous, orange-red, margin crenate to entire, apex emarginate, inner tepals 3, free, flabellate, 2.7–5.1 × 3.3–4.3 mm. Ovary trilocular with entire placentae, ovuliferous throughout; styles multifid, 2.4–5.1 mm long; stigmatic surface covering only the apices of the styles. Capsule turbinate, trilocular, with three lateral horns, with a few hairs at the base,7.6–12.9 × 14.1–30.5 mm, beak of the horns 4.1–9.6 mm long.; apical column ca. 2.9 mm long </div><div></div><h3 data-label="135091" class="ltx_title_subsubsection">Habitat and distribution </h3><div>on the border between the Colombian departments of Valle del Cauca and Chocó, in cloud forest, between 1700–2500 m. </div><div></div><h3 data-label="968916" class="ltx_title_subsubsection">Notes</h3><div>It is possible to separate <i>B. suaviola</i> from the closely related species based on both floral and vegetative characters. Because of the number of stamens (4) and the two series of tepals in the pistillate and staminate flowers this species is likely most related with <i>B. kalbreyerii</i>. However, the strigose abaxial leaf surface, and the leaf shape (oblong to elliptic versus ovate). </div><div></div><h3 data-label="878966" class="ltx_title_subsubsection">Additional specimens examined </h3><div>COLOMBIA.<b> Chocó</b>: highway Ansermanuevo–San José del Palmar, Alto del Galápago, 2050–2100 m, 25 Aug 1976, <i>Forero, E. 2076</i> (COL, MO); <i>Ibiden</i>,18 Feb 1977, <i>Forero, E. 2886</i> (COL); Municipio San José del Palmar, 1550 m, 14 Nov 1978, <i>Lozano, G. 3058</i> (COL); Kilometer 55 on the highway Ansermanuevo-San José del Palmar, 1700–1950, 19 Mar 1980, <i>Lozano, G. 3186</i> (COL); Municipio San José del Palmar, vereda El Tabor, 1450–1550 m, <i>Diaz, S. 3657</i> (COL, MO); Municipio San José del Palmar, vereda Corcovados, finca La esperanza, 1950 m, <i>Díaz, S. 3684</i> (COL, MO); <b>Valle del Cauca</b>: Municipio El Cairo, vereda El Brillante, natural reserve Cerro El Inglés, along the trail from La Fontana to Los Santicos, 2217–2328 m, 26 Sep2013, Jara, A.2397 (ANDES); <i>ibiden</i>, <i>Jara, A.2398</i> (ANDES); <i>ibiden</i>, <i>Jara, A.2399</i> (COL). <i>ibiden</i>, Jara, A.2407 (ANDES). <i>ibiden</i>, <i>Jara, A.2408</i> (UDBC); <i>Ibiden</i>, <i>Jara, A.2409</i> (HUA); <i>ibiden</i>, <i>Jara, A.2410</i> (ANDES). <i>ibiden</i>, Jara, A.2411 (UPTC); <i>ibiden</i>, 27 Sep 2013, <i>Jara, A.2447</i> (ANDES, CUVC); Municipio El Cairo, corregimiento Boquerón, vereda Las Amarillas, Serrania de Los Paraguas, ca. 4º45'N,76º20'W, along road to and beyond Cerro El Inglés, 17–23 km of El Cairo, 1750–2050 m, 13 May 1988, <i>Luteyn, J.12265</i> (CUVC); Municipio El Cairo, Cerro del Inglés, 2340 m, 28 Dec 1956, <i>Silverstone-Sopkin, F. 2671</i> (CUVC); <i>Ibiden</i>, 30 Dec 1986, <i>Silverstone-Sopkin, F. 2805 </i>(CUVC); <i>Ibiden</i>, 31dec 1986, <i>Silverstone-Sopkin, F. 2822 </i>(CUVC).</div><div></div><div><b>28.<i> Begonia toledana</i></b><i> </i>L.B. Sm. & B.G. Schub. (1946: 26). TYPE. COLOMBIA. <b>Norte de Santander</b>: Loso and vicinity, north of Toledo, 2200–2400 m, 6–7 March 1927, <i>E.P. Killip & A.C. Smith 20359</i> (holotype GH!, isotypes NY!, US!).</div><div><b>28a</b>. <b><i>Begonia toledana</i></b> var. <b><i>toledana</i></b> L.B. Sm. & B.G. Schub. </div><div>Caulescent herb, to 60–90 cm high. <i>Stem </i>erect; internodes glabrous, puberulous in the younger parts.<i> Stipules</i> sub-persistent, oblong, 8 – 12 × 1.9–2.9 mm, apex obtuse with terminal seta, light green, glabrous, margin entire. <i>Leaves</i> oblique respect to the petiole; petiole 1–4 cm long, pubescent; blade asymmetric, ovate, 5–7 × 2–3.6 cm, membranous, apex acuminate, base cordate, margin crenate-serrate, ciliate, upper surface sparsely puberulous, lower surface puberulous on veins, veins palmate-pinnate, with <span class="au-background-blue" data-background="true">X–X</span> secondary on the larger side, <span class="au-background-blue" data-background="true">X–X</span> secondary veins on the smaller side. <i>Inflorescences</i> bisexual and male; the bisexual axillary, erect, cincinum, bearing up to 3 staminate flowers and 1 pistillate flowers, protandrous; peduncle to 36 mm long, puberulus, bracts deciduous, elliptic, to 7 mm long, glabrous, apex obtuse, margin entire. <i>Male flowers</i>: pedicels to 10–12 mm long, glabrous; tepals 4, spreading, outer 2 subcordate, 8.7–10 × 9.6–11.4 mm, apex rounded, white, glabrous, margin entire, inner 2 narrow elliptic, ca. 7 × 3.2 mm, apex obtuse, white, glabrous, margin entire; stamens ca. 30, spreading, yellow, filaments <1 mm long, free, anthers linear-oblong, ca. 3.5 × 0.6 mm, lateral dehiscence, connectives produced. <i>Female flowers</i>: pedicels to 12 mm long; bracteoles likely absent; tepals 5, subequal, spreading, ca. 15.4 × 10.8 mm, apex rounded, white, glabrous, margin entire; ovary turbinate, ca. 10.2 × 12.7 mm, pubescent, equally 3-horned, horns oblong slightly upward curved, ca. 3.5 × 1 mm, placenta branches bifid; styles fusion toward the base, ca. 7 mm long, ca. 3-times divided, stigmatic papillae restricted to the apex. <i>Fruit</i> unknown.</div><h3 data-label="645840" class="ltx_title_subsubsection"> Habitat and distribution </h3><div></div><h3 data-label="589735" class="ltx_title_subsubsection">Notes</h3><div></div><h3 data-label="173145" class="ltx_title_subsubsection"> Additional specimens examined </h3><div>Only known from the type specimen.</div><div></div><div><b>28b. <i>Begonia toledana</i> </b>var. <b><i>erubescens</i></b> L.B. Sm. & B.G. Schub.(1946: 28). TYPE. COLOMBIA. Santander. </div><div>Herb, to 50 cm high. <i>Stem </i>erect; internodes to <span class="au-background-blue" data-background="true">X cm long</span>, glabrous.<i> Stipules</i> deciduous, oblong, ca. 3.8 × 1.6 mm, apex obtuse, glabrous, margin entire. <i>Leaves</i> oblique respect to the petiole, strongly asymmetric; petiole 9.5–17 mm long, glabrous; blade strongly asymmetric, ovate, 39.3 × 16 mm, apex long acuminate, base cordate, one side auriculate, rounded on the other side, margin lobulate-serrate, ciliate, upper surface dark green, sparsely setose, lower surface pale green to purplish, glabrous, veins palmate-pinnate, with 7–8 secondary on the larger side, 5–6 secondary veins on the smaller side. <i>Inflorescences</i> bisexual, axillary, erect, cicinum or dichacium with cincinum at the apex, <span class="au-background-blue" data-background="true"> bearing up to X staminate flowers and X pistillate flowers, </span>protrandrous; peduncle to 4.3 cm long, green, glabrous, bracts decidupus, oblong, ca. 2.3 × 1 mm, pale green, glabrous, obtuse, margin entire. <i>Male flowers</i>: pedicels to 8.6 mm long, glabrous; tepals 4, spreading, outer 2 ovate, 6.2–7.1 × 3.7–5.6 mm, apex rounded, base cordate, white to pinkish, <span class="au-background-blue" data-background="true">glabrous</span>, margin entire, inner 2 narrow oblong, <span class="au-background-blue" data-background="true"> X–X × X–X</span> mm, apex <span class="au-background-blue" data-background="true">rounded</span>, base shape, colour, indumentum, margin entirety, ciliation; stamens X–X, projecting/spreading, colour, f<span class="au-background-blue" data-background="true">ilaments X–X mm long, free/fusion, anthers shape, X–X × X–X mm long, dehiscence, connectives extension, symmetrically/asymmetrically fixation. </span><i><span class="au-background-blue" data-background="true">Female flowers</span></i><span class="au-background-blue" data-background="true">: pedicels to X mm long; bracteoles X, position, shape, X–X × X–X mm, apex shape, opacity, colour, indumentum, margin entirety, ciliation; tepals X, subequal/equal, persistence in fruit, projecting/spreading, the largest shape, X–X × X–X mm, apex shape, colour, indumentum, margin entirety, ciliation, the smallest shape, X–X × X–X mm, apex shape, colour, indumentum, margin entirety, ciliation; ovary body shape, X–X × X–X mm, colour, indumentum, equally/unequally X -winged, wings shape, largest X–X × X–X mm, smallest X–X × X–X mm; X-locular, placentation axillary/parietal, branches entire/divided, bearing ovules on both surfaces; styles X, colour, fusion, X–X mm long, X-lobed/X-divided, stigmatic papillae placement. </span><i><span class="au-background-blue" data-background="true">Fruiting pedicel</span></i><span class="au-background-blue" data-background="true"> to X mm long. </span><i><span class="au-background-blue" data-background="true">Fruit</span></i><span class="au-background-blue" data-background="true"> a capsule, body shape, to X × X mm, drying colour, wings same shape as in ovary, expanding to X mm tall. </span><i><span class="au-background-blue" data-background="true">Seeds</span></i><span class="au-background-blue" data-background="true"> not examined.</span></div><h3 data-label="787892" class="ltx_title_subsubsection"> </h3><h3 data-label="787892" class="ltx_title_subsubsection">Habitat and distribution </h3><div></div><h3 data-label="303823" class="ltx_title_subsubsection"> Notes </h3><div></div><h3 data-label="628961" class="ltx_title_subsubsection"> Additional specimens examined </h3><div></div><div><b>29. <i>Begonia trianae </i></b>(A. DC.) Warb. (1894: 146). <i>Casparya trianae</i> A. DC. (1859: 117). Type:–COLOMBIA. Provincia de Bogotá, Ubalá, 1750 m, 1855, <i>J.J. Triana 5048</i> (5153) (holotype BM!, isotype COL!).</div><div> Herb, to 1 m high. <i>Stem </i>erect; <span class="au-background-blue" data-background="true">internodes to X cm </span>long, pubescent.<i> Stipules</i> persistent, ovate or reniform, 8–10 <span class="au-background-blue" data-background="true">× X–X</span> mm, apex obtuse, green, pubescent, margin entire. <i>Leaves</i> oblique respect to the petiole, strongly asymmetric; petiole 12–35 mm long, <span class="au-background-blue" data-background="true">reddish</span>, pubescent; blade ovate, to 2–5 × 3–10 cm, coriaceous, apex acute, base cordate, rounded on side and auriculate on the other side, margin crenate-serrate, ciliate, upper surface green, desely pubescent, lower surface pale green, pubescent, veins pinnate/palmate, with <span class="au-background-blue" data-background="true">X–X</span> secondary on the larger side,<span class="au-background-blue" data-background="true"> X–X</span> secondary veins on the smaller side. <i>Inflorescences</i> unisexual in monoecious plants, axillary, the male inflorescence erect, cincinus, bearing up to 4 flowers; peduncle to 2.5–6 cm long, pale green, pubescent, <span class="au-background-blue" data-background="true">bracts persistence, shape, X–X × X–X mm, opacity, colour, indumentum, apex shape, margin entirety, ciliation</span>. <i>Female inflorescence</i> solitary flowers.<i> Male flowers</i>: pedicels to <span class="au-background-blue" data-background="true">X</span> mm long, pubescent; tepals 4, spreading, outer 2 suborbicular, <span class="au-background-blue" data-background="true">X–X </span>× 6–8 mm, apex rounded, base <span class="au-background-blue" data-background="true">shape</span>, white or light green, pubescent ouside, margin <span class="au-background-blue" data-background="true">entirety</span>, <span class="au-background-blue" data-background="true">ciliation</span>, inner 2 narrow obovate, <span class="au-background-blue" data-background="true">X–X × X–X</span> mm, apex <span class="au-background-blue" data-background="true">obtuse</span>, white, glabrous, margin entire; stamens ca. 30, spreading, yellow, filaments X–X mm long, free, anthers oblong, <span class="au-background-blue" data-background="true">X–X × X–X</span> mm long, connectives few produced, obtuse. <i>Female flowers</i>: pedicels to <span class="au-background-blue" data-background="true">X</span> mm long; bracteoles <span class="au-background-blue" data-background="true">3</span>, elliptic, ca. 8 mm long, apex acute, <span class="au-background-blue" data-background="true">opacity, colour, indumentum, margin entirety, ciliation</span>; tepals 5, subequal, spreading, 6–8 mm long, apex acute, white, <span class="au-background-blue" data-background="true">pubescent outside</span>, <span class="au-background-blue" data-background="true">margin entirety, ciliation, the smallest shape, X–X × X–X mm, apex shape, colour, indumentum, margin entirety, ciliation</span>; ovary broadly turbinate, <span class="au-background-blue" data-background="true">X–X × X–X</span> mm, colour, truncate at apex, pubescent, equally 3 -winged, wings ovate, <span class="au-background-blue" data-background="true">X–X × X–X</span> mm; 3-locular, placenta branches bilamellate, bearing ovules on both surfaces; styles <span class="au-background-blue" data-background="true">3</span>, white, fused toward the base, <span class="au-background-blue" data-background="true">X–X mm </span>long, multifid, stigmatic papillae restricted to the apex. <i>Fruiting pedicel</i> to <span class="au-background-blue" data-background="true">X</span> mm long. <i>Fruit</i> a capsule, body similar to the ovary, to <span class="au-background-blue" data-background="true">X × X</span> mm, drying marron, wings same shape as in ovary, expanding to X mm tall, column absent.</div><div></div><h3 data-label="379640" class="ltx_title_subsubsection">Habitat and distribution</h3><div>Endemic to the eastern basin of the Eastern Colombian Cordillera, in the department of Cundinamarca, between 1750 and 2500 m. </div><div></div><h3 data-label="175324" class="ltx_title_subsubsection">Taxonomic notes</h3><div>Mainly because the form of the fruits</div><h3 data-label="117216" class="ltx_title_subsubsection"></h3><h3 data-label="117216" class="ltx_title_subsubsection">Additional specimens examined </h3><div>COLOMBIA. <b>Cundinamarca</b>: Paramo de Guasca, eastern slope, Quebrada de Juiquin, Cordillera Oriental, alt. 2500 m., Aug. 27, 1941, J. <i>Cuatrecasas & R. Jaramillo 11990 </i>(COL, US). (OJO INLCUIR EL DE NORTE DE SANTANDER)</div><div></div><div><b>30. <i>Begonia umbellata</i> </b>Kunth. (1825: 143). Type: —COLOMBIA: Andes de Quindio, 1841, <i>M.A. Bonplant s.n.</i> (holotype P!, isotypes B!, P!).</div><div>Herb, to 1.5 m high. <i>Stem </i>initially erect and becoming prostrate; internodes to <span class="au-background-blue" data-background="true">X</span> cm long, reddish in the younger parts, glabrous.<i> Stipules</i> tardily deciduous, oblong-elliptic, 10–18 × 5.4–8.6 mm, apex obtuse, brown, soon glabrous, margin entire. <i>Leaves</i> strongly asymmetric, oblique or straight respect to the petiole; petiole 1–5 cm long, reddish, hirtellous to glabrous; lamina ovate, to 5–7 × 2.5–5 cm, coriaceous, apex acuminate, base cordate, one lobe much longer than the other, margin densely serrate, ciliate, upper surface sparsely setose above, lower surface hirtellous on the veins, veins palmate-pinnate, with 7–8 secondary on the larger side, 4–6 secondary veins on the smaller side. <i>Inflorescences</i> unisexual in monoesious plants, axillary, the male inflorescence pendant, cime umbel-like, bearing up to 20 flowers; peduncle 4–7 cm long, reddish, glabrous, bracts deciduous, elliptic to obovate, 8–12 × 6–8 mm, membranous, brown, indumentum, apex truncate, ciliate, margin entire. <i>Male flowers</i>: pedicels very slender, to 2 cm long, glabrous; tepals 4, projecting, outer 2 elliptic, 12–20 × 5–7.5 mm, apex acute, red, pilose outside, margin entire, inner 2 obovate, 13–14× 7–8 mm, apex serrate, pale red, glabrous, margin entire; stamens up to 20, projecting, pink, filaments unequal, 2–10 mm long, fused at the base, anthers ovoid, 1.8–2.2 × 1–1.2 mm, dehiscence introrse, connectives no produced. Female inflorescence, solitary flowers, peduncle to 1.5 cm, glabrous. <i>Female flowers</i>: pedicels to 38 mm long; bracteoles 2, suborbicular, ca. 12 × 9 mm, apex truncate, ciliate, red, glabrous, margin entire; tepals 6, subequal, projecting, the outer 3 thicker and narrower, 23–28 × 4.7–6.4 mm, apex lacerate-serrate, red, sparcely pubescent, margin entirety, inner 3 lanceolate, 21–23 × 7.8–9.4 mm, apex serrate, pale red, glabrous, margin entire; ovary turbinate, 10.3–12.3 × 10–12.5 mm, green and reddish, pubescent, unequally 3-horned, horns falcate, and curved upward, placenta branches bifid, bearing ovules on both surfaces; styles 3 or 4 main branches fused to the base, yellowish, 10.5–14.3 mm long, each branch dividing near to the base 1-2 times, stigmatic papillae linear. <i>Fruiting pedicel</i> to 38 mm long. <i>Fruit</i> a capsule, like the ovary, to 22 × 18 mm, horns same shape as in ovary; column 4.3–7 mm long.</div><h3 data-label="729145" class="ltx_title_subsubsection"></h3><h3 data-label="729145" class="ltx_title_subsubsection">Habitat and distribution </h3><div></div><h3 data-label="155766" class="ltx_title_subsubsection"> Taxonomic notes </h3><h3></h3><h3>Additional specimens examined </h3><div></div><div><b>31.<i> Begonia ursina</i></b><i> </i>L.B. Sm. & B.G. Schub. (1946: 28). Type:—COLOMBIA. <b>Guajira</b> (formely Magdalena): Cerro Pintado, Sierra de Perijá, 3100 m, 3–6 July 1942, <i>M.A. Carriker, Jr. 25</i> (holotype US). </div><div>Herb to 1 m. Internodes 15−30 mm long, up to 5.5 mm at cross, reddish, pubescent, nodes thickened. <i>Stipules</i> deciduous,11.6−13.7.3 x 3.8−5.6 mm, ovate, pubescent, green, coriaceous, margin entire, apex obtuse. Petioles (3)5.3−13.2 cm, hirsutous. Leaves straight, 4.9−9.5 x 2.2−3.6 mm, elliptic or oblong, asymmetric, coriaceous; base unequal, rounded to acute on one side and auriculate on the other side; apex short acuminate; margin crenate-serrulate; venation pinnate, with 6-8 major lateral veins on each side. Inflorescences unisexual in monoecious plants; <i>male inflorescences</i> bearing to 3 flowers, peduncle to 28 mm, densely pubescent, bracts desidupus, not seen. <i>Male flowers</i>: pedicels to 11.2 mm long, hirsutous; tepals 4 (rarely 6), spreading, outer 2 ovate, 10–18.3 × 6.4–12.2 mm, apex obtuse, white, hirsute outside, margin entire, setulose, inner 2 (rarely 3) obovate, 7.7–10 × 3.4–4.8 mm, apex rounded, white, escencially glabrous, with some hairs at the base outside, margin entire; stamens ca. 40, spreading, yellow, filaments 1.3–1.8 mm long, free, anthers linear-oblong, 1.9–3.5 × 0.7–1 mm, dehiscence lateral, connectives short produced, ca. 0.3 mm, obtuse. <i>Female flowers</i>: pedicels to 29 mm long; bracteoles 2, opposite, elliptic, ca. 11 × 5 mm, apex obtuse, green, pubescent, margin entire, ciliate; tepals 6, subequal, spreading, the largest shape, 13.3–23 × 7.9–15.6 mm, apex obtuse or truncate, serrulate, white, pubescent, margin serrulate toward the apex, ciliate, the smallest shape, 11.5–15.8 × 4.7–10.3 mm, apex rounded, white, pubescent, margin ciliate; ovary body turbinate, ca. 6.9 × 20.7 mm, green, hirsute, equally 3-horned, horns lanceolate; placenta branches bifid, bearing ovules on both surfaces; styles multifidi, 5–5.5 mm long, apex lobulated, stigmatic papillae restricted to the apical lobules. <i>Fruiting pedicel</i> to 44 mm long. <i>Fruit</i> a capsule, body turbinate, to 13 × 29 mm, horns same shape as in ovary; column absent.</div>