Inconsistent invocation of handler interceptors on using kekkonen operations

[metametadata]

See https://github.com/metosin/kekkonen-sample/:

1. Add a log into security/require-roles meta handler:

(defn require-roles [required]
  (fn [context]
    (println ">>>> require-roles") ; <----------
    (let [roles (-> context :user :roles)]
      (if (seq (set/intersection roles required))
        context))))

2. Run app: $ lein repl, then (reset).
3. Open Swagger UI in browser (http://localhost:3000)
4. Try out kekkonen operations (http://localhost:3000/index.html#/kekkonen).
5. See logs in REPL.

Actual:
Visiting Swagger UI hits the interceptor.
GET /kekkonen/handlers hits the interceptor.
GET /kekkonen/handler?kekkonen.action=api.admin%2Freset-counter! does not hit the interceptor.

Expected:
Both GET /kekkonen/handler and GET /kekkonen/handlers should consistently hit the handler's interceptors?